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There are 2 fourteen-letter words ending with CRONUCLEUSES

MACRONUCLEUSESMACRONUCLEUS n. a relatively large densely staining nucleus of most ciliate protozoans that is derived from micronuclei and controls various nonreproductive functions.
MICRONUCLEUSESMICRONUCLEUS n. a minute nucleus.

Scrabble words — in black are valid world wide — in RED are not valid in North America — in GREEN are valid only in North America.
Definitions are short excerpt from the 1Word.ws.

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