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There are 4 nine-letter words beginning with SUBSTR

SUBSTRACTsubstract v. (Obsolete) To subtract.
SUBSTRACT v. to subtract; to withdraw, also SUBTRACT.
SUBSTRATAsubstrata n. Plural of substratum.
SUBSTRATUM n. that which underlies something, as a layer of earth lying under another, also SUBSTRATE.
SUBSTRATEsubstrate n. An underlying layer; a substratum.
substrate n. The substance lining the bottom edge of an enclosure.
substrate n. (Biochemistry) A substance acted upon, as by an enzyme.
SUBSTRUCTsubstruct v. To build beneath something; to lay as the foundation.
SUBSTRUCT v. to build beneath; to lay as a foundation.

Scrabble words — in black are valid world wide — in RED are not valid in North America — in GREEN are valid only in North America.
Definitions are short excerpt from the WikWik.org and 1Word.ws.

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