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There are 6 fifteen-letter words containing 4A, L, O and T

ACANTHOCEPHALANacanthocephalan n. Any of parasitic intestinal worms comprising the phylum Acanthocephala.
acanthocephalan adj. In a manner similar to Acanthocephala.
ACANTHOCEPHALAN adj. belonging to the Acanthocephala, a division of parasitic worms with spiny proboscis.
PALAEOBOTANICALpalaeobotanical adj. Alternative spelling of paleobotanical.
PALAEOBOTANICAL adj. relating to palaeobotany, the study of fossil plants, also PALAEOBOTANIC.
PALATALISATIONSpalatalisations n. Plural of palatalisation.
PALATALISATION n. the process of palatalising, also PALATALIZATION.
PALATALIZATIONSpalatalizations n. Plural of palatalization.
PALATALIZATION n. the process of palatalizing, also PALATALISATION.
PHANTASMAGORIALphantasmagorial adj. Alternative form of phantasmagoric.
PHANTASMAGORIAL adj. of or like a phantasmagoria, also PHANTASMAGORIC.
TARSOMETATARSALtarsometatarsal adj. (Anatomy) Of or pertaining to both the tarsus and metatarsus.
tarsometatarsal adj. (Anatomy) Of or pertaining to the tarsometatarsus.
TARSOMETATARSAL adj. relating to the tarsometatarsus, a bird's shankbone.

Scrabble words — in black are valid world wide — in RED are not valid in North America — in GREEN are valid only in North America.
Definitions are short excerpt from the WikWik.org and 1Word.ws.

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